Tooth development is the complex process by which teeth form from embryonic cells, grow, and erupt into the mouth.. For human teeth to have a healthy oral environment, enamel, dentin, cementum, and the periodontium must all develop during appropriate stages of fetal development. Primary teeth start to form between the sixth and eighth weeks in utero, and permanent teeth begin to form in the twentieth week in utero.

 Overview

The tooth bud (sometimes called the tooth germ) is an aggregation of cells that eventually forms a tooth.These cells are derived from the ectoderm of the first branchial arch and the ectomesenchyme of the neural crest.The tooth bud is organized into three parts: the enamel organ, the dental papilla and the dental follicle.

The enamel organ is composed of the outer enamel epithelium, inner enamel epithelium, stellate reticulum and stratum intermedium.These cells give rise to ameloblasts, which produce enamel and the reduced enamel epithelium. The location where the outer enamel epithelium and inner enamel epithelium join is called the cervical loop. The growth of cervical loop cells into the deeper tissues forms Hertwig's Epithelial Root Sheath, which determines the root shape of the tooth.

The dental papilla contains cells that develop into odontoblasts, which are dentin-forming cells. Additionally, the junction between the dental papilla and inner enamel epithelium determines the crown shape of a tooth. Mesenchymal cells within the dental papilla are responsible for formation of tooth pulp.

The dental follicle gives rise to three important entities: cementoblasts, osteoblasts, and fibroblasts. Cementoblasts form the cementum of a tooth. Osteoblasts give rise to the alveolar bone around the roots of teeth. Fibroblasts develop the periodontal ligaments which connect teeth to the alveolar bone through cementum.

MANDIBULAR SECOND BICUSPID

Facial: From this aspect, the tooth somewhat resembles the first, but the buccal cusp is less pronounced. The tooth is larger than the first.

Lingual: Two significant variations are seen in this view. The most common is the three-cusp form which has two lingual cusps. The mesial of those is the larger of the two. The other form is the two-cusp for with a single lingual cusp. In that variant, the lingual cusp tip is shifted to the mesial.

Proximal: The buccal cusp is shorter than the first. The lingual cusp (or cusps) are much better developed than the first and give the lingual a full, well-developed profile.

Occlusal: The two or three cusp versions become clearly evident. In the three-cusp version, the developmental grooves present a distinctive 'Y' shape and have a central pit. In the two cusp version, a single developmental groove crosses the transverse ridge from mesial to distal

Contact Points; Height of Curvature: From the facial, the mesial contact is more occlusal than the distal contact.The distal marginal ridge is lower than the mesial marginal ridge

Root Surface:-The root of the tooth is single, that is usually larger than that of the first premolar  

the lower second premolar is larger than the first, while the upper first premolar is just slightly larger than the upper second

There may be one or two lingual cusps

MAXILLARY SECOND MOLAR

The second molars are often called 12-year molars because they erupt when a child is about 12 years

Facial: The crown is shorter occluso-cervically and narrower mesiodistally whe compared to the first molar. The distobuccal cusp is visibly smaller than the mesiobuccal cusp. The two buccal roots are more nearly parallel. The roots are more parallel; the apex of the mesial root is on line with the with the buccal developmental groove. Mesial and distal roots tend to be about the same length.

Lingual: The distolingual cusp is smaller than the mesiolingual cusp. The Carabelli trait is absent.

Proximal: The crown is shorter than the first molar and the palatal root has less diverence. The roots tend to remain within the crown profile.

Occlusal: The distolingual cusp is smaller on the second than on the first molar. When it is much reduced in size, the crown outline is described as 'heart-shaped.' The Carabelli trait is usually absent. The order of cusp size, largest to smallest, is the same as the first but is more exaggerated: mesiolingual, mesiobuccal, distobuccal, and distolingual.

Contact Points; Height of Curvature: Both mesial and distal contacts tend to be centered buccolingually below the marginal ridges. Since themolars become shorter, moving from first to this molar, the contacts tend to appear more toward the center of the proximal surfaces.

Roots: There are three roots, two buccal and one lingual. The roots are less divergent than the first with their apices usually falling within the crown profile. The buccal roots tend to incline to the distal.

Note: The distolingual cusp is the most variable feature of this tooth. When it is large, the occlusal is somewhat rhomboidal; when reduced in size the crown is described as triangual or 'heart-shaped.' At times, the root may be fused.

MAXILLARY CENTRAL INCISORS

Viewed mesially or distally, a maxillary central incisor looks like a wedge, with the point of the wedge at the incisal (cutting) edge of the tooth.

Facial Surface- The mesial margin is nearly straight and meets the incisal edge at almost a 90° angle, but the distal margin meets the incisal edge in a curve. The incisal edge is straight, but the cervical margin is curved like a half moon. Two developmental grooves are on the facial surface.

Lingual Surface:- The lingual aspect presents a distinctive lingual fossa that is bordered by mesial and distal marginal ridges, the incisal edge, and the prominent cingulum at the gingival. Sometimes a deep pit, the lingual pit, is found in conjunction with a cingulum.

Incisal: The crown is roughly triangular in outline; the incisal edge is nearly a straight line, though slightly crescent shaped

Contact Points: The mesial contact point is just about at the incisal, owing to the very sharp mesial incisal angle. The distal contact point is located at the junction of the incisal third and the middle third.

Root Surface:-As with all anterior teeth, the root of the maxillary central incisor is single. This root is from one and one-fourth to one and one-half times the length of the crown. Usually, the apex of the root is inclined slightly distally.