As root and cementum formation begin, bone is created in the adjacent area. Throughout the body, cells that form bone are called osteoblasts. In the case of alveolar bone, these osteoblast cells form from the dental follicle. Similar to the formation of primary cementum, collagen fibers are created on the surface nearest the tooth, and they remain there until attaching to periodontal ligaments.

Like any other bone in the human body, alveolar bone is modified throughout life. Osteoblasts create bone and osteoclasts destroy it, especially if force is placed on a tooth. As is the case when movement of teeth is attempted through orthodontics, an area of bone under compressive force from a tooth moving toward it has a high osteoclast level, resulting in bone resorption. An area of bone receiving tension from periodontal ligaments attached to a tooth moving away from it has a high number of osteoblasts, resulting in bone formation.

MAXILLARY SECOND BICUSPID

smaller in dimensions. The cusps are not as sharp as the maxillary first bicuspid and have only one root.

Facial: This tooth closely resembles the maxillary first premolar but is a less defined copy of its companion to the mesial. The buccal cusp is shorter, less pointed, and more rounded than the first.

Lingual: Again, this tooth resembles the first. The lingual cusp, however, is more nearly as large as the buccal cusp.

Proximal: Mesial and distal surfaces are rounded. The mesial developmental depression and mesial marginal ridge are not present on the second premolar.

Occlusal: The crown outline is rounded, ovoid, and is less clearly defined than is the first.

Contact Points; When viewed from the facial, the distal contact area is located more cervically than is the mesial contact area.

Dentinogenesis

Dentin formation, known as dentinogenesis, is the first identifiable feature in the crown stage of tooth development. The formation of dentin must always occur before the formation of enamel. The different stages of dentin formation result in different types of dentin: mantle dentin, primary dentin, secondary dentin, and tertiary dentin.

Odontoblasts, the dentin-forming cells, differentiate from cells of the dental papilla. They begin secreting an organic matrix around the area directly adjacent to the inner enamel epithelium, closest to the area of the future cusp of a tooth. The organic matrix contains collagen fibers with large diameters (0.1-0.2 μm in diameter). The odontoblasts begin to move toward the center of the tooth, forming an extension called the odontoblast process. Thus, dentin formation proceeds toward the inside of the tooth. The odontoblast process causes the secretion of hydroxyapatite crystals and mineralization of the matrix. This area of mineralization is known as mantle dentin and is a layer usually about 150 μm thick.

Whereas mantle dentin forms from the preexisting ground substance of the dental papilla, primary dentin forms through a different process. Odontoblasts increase in size, eliminating the availability of any extracellular resources to contribute to an organic matrix for mineralization. Additionally, the larger odontoblasts cause collagen to be secreted in smaller amounts, which results in more tightly arranged, heterogenous nucleation that is used for mineralization. Other materials (such as lipids, phosphoproteins, and phospholipids) are also secreted.

Secondary dentin is formed after root formation is finished and occurs at a much slower rate. It is not formed at a uniform rate along the tooth, but instead forms faster along sections closer to the crown of a tooth. This development continues throughout life and accounts for the smaller areas of pulp found in older individuals. Tertiary dentin, also known as reparative dentin, forms in reaction to stimuli, such as attrition or dental caries.

The dentin in the root of a tooth forms only after the presence of Hertwig's epithelial root sheath (HERS), near the cervical loop of the enamel organ. Root dentin is considered different than dentin found in the crown of the tooth (known as coronal dentin) because of the different orientation of collagen fibers, the decrease of phosphoryn levels, and the less amount of mineralization.

Deciduous dentition period.

-The deciduous teeth start to erupt at the age of six months and the deciduous dentition is complete by the age of approximately two and one half years of age.

-The jaws continue to increase in size at all points until about age one year.

-After this, growth of the arches is lengthening of the arches at their posterior (distal) ends. Also, there is slightly more forward growth of the mandible than the maxilla.

1. Many early developmental events take place.

-The tooth buds anticipate the ultimate occlusal pattern.

-Mandibular teeth tend to erupt first. The pattern for the deciduous incisors is usually in this distinctive order:

(1) mandibular central

(2) maxillary central incisors

(3) then all four lateral incisors.

-By one year, the deciduous molars begin to erupt.

-The eruption pattern for the deciduous dentition as a whole is:

(1) central incisor

(2) lateral incisor

(3) deciduous first molar

(4) then the canine

(5) then finally the second molar.

-Eruption times can be variable.

2. Occlusal changes in the deciduous dentition.

-The overjet tends to diminish with age. Wear and mandibular growth are a factor in this process.

-The overbite often diminishes with the teeth being worn to a flat plane occlusion.

-Spacing of the incisors in anticipation of the soon-to-erupt permanent incisors appears late. Permanent anterior teeth (incisors and canines) are wider mesiodistally than deciduous anterior teeth. In contrast, the deciduous molar are wider mesiodistally that the premolars that later replace them.

-Primate spaces occur in about 50% of children. They appear in the deciduous dentition. The spaces appear between the upper lateral incisor and the upper canine. They also appear between the lower canine and the deciduous first molar.

HISTOLOGIC CHANGES OF THE PULP

Regressive changes

Pulp decreases in size by the deposition of dentin.
This can be caused by age, attrition, abrasion, operative procedures, etc.
Cellular organelles decrease in number.

Fibrous changes

They are more obvious in injury rather than aging. Occasionally, scarring may also be apparent.

Pulpal stones or denticles

They can be: a)free, b)attached and/or c)embedded. Also they are devided in two groups: true or false. The true stones (denticles) contain dentinal tubules. The false predominate over the the true and are characterized by concentric layers of calcified material.

Diffuse calcifications

Calcified deposits along the collagen fiber bundles or blood vessels may be observed. They are more often in the root canal portion than the coronal area.

Histology of the Cementum

Cementum is a hard connective tissue that derives from ectomesenchyme.

Embryologically, there are two types of cementum:
Primary cementum: It is acellular and develops slowly as the tooth erupts. It covers the coronal 2/3 of the root and consists of intrinsic and extrinsic fibers (PDL).
Secondary cementum: It is formed after the tooth is in occlusion and consists of extrinsic and intrinsic (they derive from cementoblasts) fibers. It covers mainly the root surface.

Functions of Cementum

It protects the dentin (occludes the dentinal tubules)
It provides attachment of the periodontal fibers
It reverses tooth resorption

Cementum is composed of 90% collagen I and III and ground substance.
50% of cementum is mineralized with hydroxyapatite. Thin at the CE junction, thicker apically.

MAXILLARY FIRST BICUSPID (PREMOLARS)

It is considered to be the typical bicuspid. (The word "bicuspid" means "having two cusps.")

Facial: The buccal surface is quite rounded and this tooth resembles the maxillary canine. The buccal cusp is long; from that cusp tip, the prominent buccal ridge descends to the cervical line of the tooth.

Lingual: The lingual cusp is smaller and the tip of that cusp is shifted toward the mesial. The lingual surface is rounded in all aspects.

Proximal: The mesial aspect of this tooth has a distinctive concavity in the cervical third that extends onto the root. It is called variously the mesial developmental depression, mesial concavity, or the 'canine fossa'--a misleading description since it is on the premolar. The distal aspect of the maxillary first permanent molar also has a developmental depression. The mesial marginal developmental groove is a distinctive feature of this tooth.

Occlusal: There are two well-defined cusps buccal and lingual. The larger cusp is the buccal; its cusp tip is located midway mesiodistally. The lingual cusp tip is shifted mesially. The occlusal outline presents a hexagonal appearance. On the mesial marginal ridge is a distinctive feature, the mesial marginal developmental groove.

Contact Points;The distal contact area is located more buccal than is the mesial contact area.

Root Surface:-The root is quite flat on the mesial and distal surfaces. In about 50 percent of maxillary first bicuspids, the root is divided in the apical third, and when it so divided, the tips of the facial and lingual roots are slender and finely tapered.