Bell stage

The bell stage is known for the histodifferentiation and morphodifferentiation that takes place. The dental organ is bell-shaped during this stage, and the majority of its cells are called stellate reticulum because of their star-shaped appearance. Cells on the periphery of the enamel organ separate into three important layers. Cuboidal cells on the periphery of the dental organ are known as outer enamel epithelium.The cells of the enamel organ adjacent to the dental papilla are known as inner enamel epithelium. The cells between the inner enamel epithelium and the stellate reticulum form a layer known as the stratum intermedium. The rim of the dental organ where the outer and inner enamel epithelium join is called the cervical loop

Other events occur during the bell stage. The dental lamina disintegrates, leaving the developing teeth completely separated from the epithelium of the oral cavity; the two will not join again until the final eruption of the tooth into the mouth

The crown of the tooth, which is influenced by the shape of the internal enamel epithelium, also takes shape during this stage. Throughout the mouth, all teeth undergo this same process; it is still uncertain why teeth form various crown shapes—for instance, incisors versus canines. There are two dominant hypotheses. The "field model" proposes there are components for each type of tooth shape found in the ectomesenchyme during tooth development. The components for particular types of teeth, such as incisors, are localized in one area and dissipate rapidly in different parts of the mouth. Thus, for example, the "incisor field" has factors that develop teeth into incisor shape, and this field is concentrated in the central incisor area, but decreases rapidly in the canine area. The other dominant hypothesis, the "clone model", proposes that the epithelium programs a group of ectomesenchymal cells to generate teeth of particular shapes. This group of cells, called a clone, coaxes the dental lamina into tooth development, causing a tooth bud to form. Growth of the dental lamina continues in an area called the "progress zone". Once the progress zone travels a certain distance from the first tooth bud, a second tooth bud will start to develop. These two models are not necessarily mutually exclusive, nor does widely accepted dental science consider them to be so: it is postulated that both models influence tooth development at different times.Other structures that may appear in a developing tooth in this stage are enamel knots, enamel cords, and enamel niche.

Classification of Cementum

1. Embryologically

Primary and secondary

2. According to cellular component

Acellular: Thin, Amorphous, First layer to seal the dentin tubules

Cellular: Thick, Better structure, Apical surface

Layers of cellular and acellular cementum alternate (randomly)

3. Based on the origin of the collagenous matrix
Extrinsic
Intrinsic
Mixed

4. Combined classification
a. Primary acellular intinsic fiber cementum
b. Primary acellualar extrinsic fiber cementum
c. Secondary cellular intrinsic fiber cementum
d. Secondary cellular mixed fiber cementum
e. Acellular afibrillar cementum

5. Depending on the location and patterning
Intermediate and mixed stratified cementum

Participating Cells

Cementoblasts

Active
Cells are round, plump with basophilic cytoplasm (rough endoplasmic reticulum)
Inactive
Cells have little cytoplasm
Cementocytes

1. Cementocyte lacuna
2. cementocyte canaliculus

Cells have fewer organelles compared to cementoblasts. They are found in lacunae and have numerous processes toward the periodontal ligament. Eventually they die due to avascularity

Cementicles

a) free
b) attached
c) embedded

CONTACT POINT.:-The point on the proximal surface where two adjacent teeth actually touch each other is called a contact point.

INTERPROXIMAL SPACE.:-The interproximal space is the area between the teeth. Part of the interproximal space is occupied by the interdental papilla. The interdental papilla is a triangular fold of gingival tissue. The part of the interproximal space not occupied is called the embrasure.

EMBRASURE. :-The embrasure occupies an area bordered by interdental papilla, the proximal surfaces of the two adjacent teeth, and the contact point (fig 4-18). If there is no contact point between the teeth, then the area between them is called a diastema instead of an embrasure.

OCCLUSAL

The occlusal surface is the broad chewing surface found on posterior teeth (bicuspids and molars).

OCCLUSION.:-Occlusion is the relationship between the occlusal surfaces of maxillary and mandibular teeth when they are in contact. Many patterns of tooth contact are possible. Part of the reason for the variety is the mandibular condyle's substantial range of movement within the temporal mandibular joint.

Malocclusion occurs when any abnormality in occlusal relationships exist in the dentition. Centric occlusion, is the centered contact position of the chewing surfaces of mandibular teeth on the chewing surface (occlusal) of the maxillary teeth.

OCCLUSAL PLANE.:-Maxillary and mandibular teeth come into centric occlusion and meet along anteroposterior and lateral curves. The anteroposterior curve is called the Curve of Spee  in which the mandibular arch forms a concave (a bowl-like upward curve). The lateral curve is called the Curve of Wilson . The composite (combination) of these curves form a line called the occlusal plane, and is created by the contact of the upper and lower teeth

VERTICAL AND HORIZONTAL OVERLAP. :-Vertical overlap is the extension of the maxillary teeth over the mandibular counterparts in a vertical direction when the dentition is in centric occlusion Horizontal overlap is the projection of maxillary teeth over antagonists (something that opposes another) in a horizontal direction.

KEY TO OCCLUSION.:-The occlusal surfaces of opposing teeth bear a definite relationship to each other. In normal jaw relations and when teeth are of normal size and in the correct position, the mesiofacial cusp of the maxillary first molar occludes in the facial groove of the mandibular first molar. This normal relationship of these two teeth is called the key to occlusion.

PERMANENT DENTITION

The permanent dentition consists of 32 teeth. Each tooth in the permanent dentition is described in this section. It should be remembered that teeth show considerable variation in size, shape, and other characteristics from one person to another. Certain teeth show a greater tendency than others to deviate from the normal. The descriptions that follow are of normal teeth.

Permanent teeth

1. The permanent teeth begin formation between birth and 3 years of age (except for the third molars)

2. The crowns of permanent teeth are completed between 4 and 8 years of age, at approximately one- half the age of eruption

The sequence for permanent development

Maxillary                     

First molar → Central incisor → Lateral incisor → First premotar → Second pmmolar  → Canine → Second molar → Third molar

Mandibular

First molar → Central incisor → Lateral incisor → Canine → First premolar → Second premolar → Second molar → Third molar

Permanent teeth emerge into the oral cavity as

                                      Maxillary                       Mandibular

Central incisor               7-8 years                        6-7 years

Lateral incisor                8-9 years                        7-8 years

Canine                           11-12 years                    9-10 years

First premolar                10-Il years                      10-12 years

Second premolar            10-12 years                  11-12 years

First molar                       6-7 years                      6-7 years

Second molar                 12-13 years                    11-13 years

Third molar                      17-21 years                    17-21 years

The roots of the permanent teeth are completed between 10 and 16 years of age, 2 to 3 years after eruption

Dentin

Composition: 70% inorganic, 20% organic, 10% water by weight and 45%, 33%, and 22% in volume respectively
Hydroxyapatite crystals and collagen type I
Physical characteristics: Harder than bone and softer than enamel
Yellow in color in normal teeth
Radiographic appearance: More radiolucent than enamel

Primary (circumpulpal) dentin: forms most of the tooth
Mantle dentin: first dentin to form; forms the outline of dentin in the adult tooth
Predentin: lines the innermost portion of dentin (faces the pulp)
Secondary dentin: after root formation dentin continues to form, continuous to primary dentin but with structural irregularities
Tertiary dentin: reactive or reparative dentin; may or may not have characteristics of primary dentin; produced in the area of an external stimulus; osteodentin

Dentin is formed by cells called odontoblasts.
These cells derive from the ectomesenchyme and produce the organic matrix that will calcify and become the dentin.
Formation of dentin initiates formation of enamel.
The formation of dentin starts during late bell-stage in the area of the future cusp.

First coronal dentin and then root dentin.

Completion of dentin does not occur until about 18 months after eruption of primary and 2-3 years after eruption of permanent teeth.

The rate of dentin development varies.

The role of the internal (inner) dental (enamel) epithelium
Cuboidal - Columnar (reverse polarization)
Ectomesenchymal cells of the dental papilla become preodontoblasts - odontoblasts
Acellular zone disappears

Histologic features of dentin
Odontoblasts
Dentinal tubules
Extend through the entire thickness of dentin
S-shaped (primary curvatures) path in the crown, less S-shaped in the root, almost straight in the cervical aspect
Secondary curvatures
Tubular microbranches
Presence of fluid
 

Intratubular dentin
Dentin in the tubule that is hypermineralized

The term peritubular dentin should not be used
 

Sclerotic dentin
Dentinal tubules that are occluded with calcified material
Most likely a physiologic response
Reduction of permeability of dentin
 

Intertubular dentin
Dentin between the tubules
 

Interglobular dentin
Areas of unmineralized or hypomineralized dentin
The defect affects mineralization and not the architecture of dentin
 

Incremental lines
Lines of von Ebner: lines associated with 5-day rythmic pattern of dentin deposition
Contour lines of Owen: Originally described by Owen they result from a coincidence of the secondary curvatures between neighboring dentinal tubules.
 

Granular Layer of Tomes
Seen only in ground sections in the root area covered by cementum
Originally, they were thought to be areas of hypomineralization
They are true spaces obtained by sections going through the looped terminal portions dentinal tubules

DE junction :Scalloped area

Enamel tissue with incremental lines of Retzius and dentin tissue with parallel, curved dentinal tubules are in contact at the irregular dentino-enamel junction. The junction often has a scalloped-shaped morphology

DC junction Dentin Cemental Junction

ERUPTION OF THE PERMANENT TOOTH

- At the time at which the deciduous tooth erupts the tooth bud for the permanent tooth has already been building up enamel and dentin.

- When the permanent tooth starts to erupt, pressure on the root of the deciduous tooth causes resorption by the osteoclasts.

- Wolff's law states that when two hard tissues exert pressure on one another the softer of the tissues will be resorbed.

- The dentin and cementum of the root of the deciduous tooth is softer than the enamel of the permanent tooth that is why the root of the deciduous tooth is resorbed.

- Most permanent teeth have erupted and have been in use for 2 years before the root is completely formed.