Cementum

Composition

a. Inorganic (50%)—calcium hydroxyapatite crystals.

b. Organic (50%)—water, proteins, and type I collagen.

c. Note: Compared to the other dental tissues, the composition of cementum is most similar to bone; however, unlike bone, cementum is avascular (i.e., no Haversian systems or other vessels are present).

Main function of cementum is to attach PDL fibers to the root surface.

Cementum is generally thickest at the root apex and in interradicular areas of multirooted 

Types of cementum

a. Acellular (primary) cementum

(1) A thin layer of cementum that surrounds the root, adjacent to the dentin.

(2) May be covered by a layer of cellular cementum, which most often occurs in the middle and apical root.

(3) It does not contain any cells.

b. Cellular (secondary) cementum

(1) A thicker, less-mineralized layer of cementum that is most prevalent along the apical root and in interradicular (furcal) areas of multirooted teeth.

(2) Contains cementocytes.

(3) Lacunae and canaliculi:

(a) Cementocytes (cementoblasts that become trapped in the extracellular matrix during cementogenesis) are observed in their entrapped spaces, known as lacunae.

(b) The processes of cementocytes extend through narrow channels called canaliculi.

(4) Microscopically, the best way to differentiate between acellular and cellular cementum is the presence of lacunae in cellular cementum.

MANDIBULAR SECOND BICUSPID

Facial: From this aspect, the tooth somewhat resembles the first, but the buccal cusp is less pronounced. The tooth is larger than the first.

Lingual: Two significant variations are seen in this view. The most common is the three-cusp form which has two lingual cusps. The mesial of those is the larger of the two. The other form is the two-cusp for with a single lingual cusp. In that variant, the lingual cusp tip is shifted to the mesial.

Proximal: The buccal cusp is shorter than the first. The lingual cusp (or cusps) are much better developed than the first and give the lingual a full, well-developed profile.

Occlusal: The two or three cusp versions become clearly evident. In the three-cusp version, the developmental grooves present a distinctive 'Y' shape and have a central pit. In the two cusp version, a single developmental groove crosses the transverse ridge from mesial to distal

Contact Points; Height of Curvature: From the facial, the mesial contact is more occlusal than the distal contact.The distal marginal ridge is lower than the mesial marginal ridge

Root Surface:-The root of the tooth is single, that is usually larger than that of the first premolar  

the lower second premolar is larger than the first, while the upper first premolar is just slightly larger than the upper second

There may be one or two lingual cusps

Soft Oral Tissues

Oral Mucosa

The oral mucosa consists mainly of two types of tissues: the oral epithelium, which consists of stratified, squamous epithelium, and the underlying connective tissue layer, known as the lamina propria.  There are three variations of oral mucosa.

A. Oral epithelium

1. Consists of stratified, squamous epithelium.

2. Four layers (Note: Cells mature as they progress from the deepest [basal] layer to the most superficial [cornified] layer) a. Basal layer (stratum germinativum or basale)

(1) A single layer of cuboidal or columnar cells overlying the lamina propria.

(2) Contains progenitor cells and thus provides cells to the epithelial layers above.

(3) Site of cell division (mitosis).

b. Prickle cell layer (stratum spinosum)

(1) Consists of several layers of larger, ovoid-shaped cells.

c. Granular layer (stratum granulosum)

(1) Cells appear larger and flattened.

(2) Granules (known as keratohyaline granules) are present in the cells.

(3) This layer is absent in nonkeratinized epithelium.

d. Cornified layer (stratum corneum, keratin, or horny layer)

(1) In keratinized epithelium:

(a) Orthokeratinized epithelium the squamous cells on the surface appear flat and contain keratin. They have no nuclei present.

(b) Parakeratinized epithelium the squamous cells appear flat and contain keratin; nuclei are present within the cells.

(2) In parakeratinized epithelium, both squamous cells without nuclei and cells with shriveled (pyknotic) nuclei are present.

(3) In nonkeratinized epithelium, the cells appear slightly flattened and contain nuclei.

B. Lamina propria

1. Consists of type I and III collagen, elastic fibers, and ground substance. It also contains many cell types, including fibroblasts, endothelial cells, immune cells, and a rich vascular and nerve supply.

2. Two layers:

a. Superficial, papillary layer

(1) Located around and between the epithelial ridges.

(2) Collagen fibers are thin and loosely arranged.

b. Reticular layer

(1) Located beneath the papillary layer.

(2) Collagen fibers are organized in thick, parallel bundles.

C. Types of oral mucosa

1. Masticatory mucosa

a. Found in areas that have to withstand compressive and shear forces.

b. Clinically, it has a rubbery, firm texture.

c. Regions: gingiva, hard palate.

2. Lining mucosa

a. Found in areas that are exposed to high levels of friction, but must also be mobile and distensible.

b. Clinically, it has a softer, more elastic texture.

c. Regions: alveolar mucosa, buccal mucosa, lips, floor of the mouth, ventral side of the tongue, and soft palate.

3. Specialized mucosa

a. Similar to masticatory mucosa, specialized mucosa is able to tolerate high compressive

and shear forces; however, it is unique in that it forms lingual papillae.

b. Region: dorsum of the tongue.

D. Submucosa

1. The connective tissue found beneath the mucosa . It contains blood vessels and nerves and may also contain fatty tissue and minor salivary glands.

2. Submucosa is not present in all regions of the oral cavity, such as attached gingiva, the tongue, and hard palate. Its presence tends to increase the mobility of the tissue overlying it.

E. Gingiva

1. The portion of oral mucosa that attaches to the teeth and alveolar bone.

2. There are two types of gingiva: attached and free gingiva. The boundary at which they meet is known as the free gingival groove .

a. Attached gingiva

(1) Directly binds to the alveolar bone and tooth.

(2) It extends from the free gingival groove to the mucogingival junction.

b. Free gingiva

(1) Coronal to the attached gingiva, it is not bound to any hard tissue.

(2) It extends from the gingival margin to the free gingival groove.

c. Together, the free and attached gingiva form the interdental papilla.

.F. Alveolar mucosa

1. The tissue just apical to the attached gingiva.

2. The alveolar mucosa and attached gingiva meet at the mucogingival junction .

G. Junctional epithelium

1. Area where the oral mucosa attaches to the tooth, forming the principal seal between the oral cavity and underlying tissues.

2. Is unique in that it consists of two basal lamina, an internal and external . The internal basal lamina, along with hemidesmosomes, comprises the attachment apparatus (the epithelial attachment). This serves to attach the epithelium directly to the tooth.

3. Histologically, it remains as immature, poorly differentiated tissue. This allows it to maintain its ability to develop hemidesmosomal attachments.

4. Has the highest rate of cell turnover of any oral mucosal tissue.

H. Interdental papilla (interdental gingiva)

1. Occupies the interproximal space between two teeth. It is formed by free and attached gingiva.

2. Functions to prevent food from entering the (interproximal) area beneath the contact point of two adjacent teeth. It therefore plays an important role in maintaining the health of the gingiva.

3. Col

a. If the interdental papilla is cross-sectioned in a buccolingual plane, it would show two peaks (buccal and lingual) with a dip between them, known as the col or interdental col. This depression occurs around the contact point of the two adjacent teeth.

b. Histologically, col epithelium is the same as junctional epithelium

Tooth eruption Theories

Tooth eruption occurs when the teeth enter the mouth and become visible. Although researchers agree that tooth eruption is a complex process, there is little agreement on the identity of the mechanism that controls eruption. Some commonly held theories that have been disproven over time include: (1) the tooth is pushed upward into the mouth by the growth of the tooth's root, (2) the tooth is pushed upward by the growth of the bone around the tooth, (3) the tooth is pushed upward by vascular pressure, and (4) the tooth is pushed upward by the cushioned hammock. The cushioned hammock theory, first proposed by Harry Sicher, was taught widely from the 1930s to the 1950s. This theory postulated that a ligament below a tooth, which Sicher observed on under a microscope on a histologic slide, was responsible for eruption. Later, the "ligament" Sicher observed was determined to be merely an artifact created in the process of preparing the slide.

The most widely held current theory is that while several forces might be involved in eruption, the periodontal ligaments provide the main impetus for the process. Theorists hypothesize that the periodontal ligaments promote eruption through the shrinking and cross-linking of their collagen fibers and the contraction of their fibroblasts.

Although tooth eruption occurs at different times for different people, a general eruption timeline exists. Typically, humans have 20 primary (baby) teeth and 32 permanent teeth. Tooth eruption has three stages. The first, known as deciduous dentition stage, occurs when only primary teeth are visible. Once the first permanent tooth erupts into the mouth, the teeth are in the mixed (or transitional) dentition. After the last primary tooth falls out of the mouth—a process known as exfoliation—the teeth are in the permanent dentition.

Primary dentition starts on the arrival of the mandibular central incisors, usually at eight months, and lasts until the first permanent molars appear in the mouth, usually at six years. The primary teeth typically erupt in the following order: (1) central incisor, (2) lateral incisor, (3) first molar, (4) canine, and (5) second molar. As a general rule, four teeth erupt for every six months of life, mandibular teeth erupt before maxillary teeth, and teeth erupt sooner in females than males. During primary dentition, the tooth buds of permanent teeth develop below the primary teeth, close to the palate or tongue.

Mixed dentition starts when the first permanent molar appears in the mouth, usually at six years, and lasts until the last primary tooth is lost, usually at eleven or twelve years. Permanent teeth in the maxilla erupt in a different order from permanent teeth on the mandible. Maxillary teeth erupt in the following order: (1) first molar (2) central incisor, (3) lateral incisor, (4) first premolar, (5) second premolar, (6) canine, (7) second molar, and (8) third molar. Mandibular teeth erupt in the following order: (1) first molar (2) central incisor, (3) lateral incisor, (4) canine, (5) first premolar, (6) second premolar, (7) second molar, and (8) third molar. Since there are no premolars in the primary dentition, the primary molars are replaced by permanent premolars. If any primary teeth are lost before permanent teeth are ready to replace them, some posterior teeth may drift forward and cause space to be lost in the mouth. This may cause crowding and/or misplacement once the permanent teeth erupt, which is usually referred to as malocclusion. Orthodontics may be required in such circumstances for an individual to achieve a straight set of teeth.

The permanent dentition begins when the last primary tooth is lost, usually at 11 to 12 years, and lasts for the rest of a person's life or until all of the teeth are lost (edentulism). During this stage, third molars (also called "wisdom teeth") are frequently extracted because of decay, pain or impactions. The main reasons for tooth loss are decay or periodontal disease.

Transient structures during tooth development

Enamel knot: Thickening of the internal dental epithelium at the center of the dental organ.
Enamel cord: Epithelial proliferation that seems to divide the dental organ in two.
 

Review the role of these two structures
Enamel niche: It is an artifact that is produced during section of the tissue. It occurs because the dental organ is a sheet of proliferating cells rather than a single strand. It looks like a concavity that contains ectomesenchyme.

Stationary Relationship

a) .Centric Relation is the most superior relationship of the condyle of the mandible to the articular fossa of the temporal bone as determined by the bones ligaments. and muscles of the temporomandibular joint; in an ideal dentition it is the same as centric occlusion.

(b) Canines may also be used to confirm the molar relationships to classify occlusion when molars are missing; a class I canine relationship shows the cusp tip of the maxillary canine facial to the mesiobuccal cusp of the first permanent molar

c) Second primary molars are used to classify the occlusion in a primary dentition

(d) In a mixed  dentition the first permanent molars will erupt into a normal occlusion if there is a terminal step between the distal  surfaces of maxillarv and mandibular second primary molars; if these surfaces are flush, a terminal plane exists and the first permanent molars will first erupt into an end-to-end relationship until there is a shifting of space or exfoliation of the second primary molar